A proxy server, also known as a "proxy" or "application level gateway", is a computer that acts as a gateway between a local network (e.g., all the computers at one company or in one building) and a larger-scale network such as the Internet. Proxy servers provide increased performance and security. In some cases, they monitor employees' use of outside resources.
A proxy server works by intercepting connections between sender and receiver. All incoming data enters through one port and is forwarded to the rest of the network via another port. By blocking direct access between two networks, proxy servers make it much more difficult for hackers to get internal addresses and details of a private network.
Some proxy servers are a group of applications or servers that block common Internet services. For example, an HTTP proxy intercepts web access, and an SMTP proxy intercepts email. A proxy server uses a network addressing scheme to present one organization-wide IP address to the Internet. The server funnels all user requests to the Internet and returns responses to the appropriate users. In addition to restricting access from outside, this mechanism can prevent inside users from reaching specific Internet resources (e.g., certain web sites). A proxy server can also be one of the components of a firewall.
Proxies may also cache web pages. Each time an internal user requests a URL from outside, a temporary copy is stored locally. The next time an internal user requests the same URL, the proxy can serve the local copy instead of retrieving the original across the network, improving performance.
Note: Do not confuse a proxy server with a NAT (Network Address Translation) device. A proxy server connects to, responds to, and receives traffic from the Internet, acting on behalf of the client computer, while a NAT device transparently changes the origination address of traffic coming through it before passing it to the Internet.
For those who understand the OSI (Open System Interconnection) model of networking, the technical difference between a proxy and a NAT is that the proxy server works on the transport layer (layer 4) or higher of the OSI model, whereas a NAT works on the network layer (layer 3).
source: http://kb.iu.edu/data/ahoo.html
related source: http://www3.rad.com/networks/1998/proxy/proxy.htm
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